Steenbok
Raphicerus campestris (Thunberg, 1811)
Photo: Doug Lee
| Afrikaans: | Steenbok |
| German: | Steinböckchen |
| French: | Steenbok |
| Swahili: | Isha / Dondor |
| isiNdebele: | Ingina |
| isiZulu: | Iqhina |
| isiXhosa: | Itshabanqa |
| seSotho: | Thiane |
| seTswana: | Phuduhudu |
| Shona: | Mhene Ndunguza / Chengu |
| Shangaan: | Shipene Ndakadsi |
| Nama: | !Aris |
IUCN Conservation Status:
Survival threat rating: Lower Risk, least concerned (LR/lc)
This graceful buck has the misfortune to be named after a brick,
perhaps because of its colour, and is the symbol of cleanliness among
the African people. It has the second widest distribution of African
antelope due to its ability to survive in marginal areas where larger,
faster and more aggressive ungulates have disappeared following human
pressure. In addition, its ability to feed almost exclusively on forbs
in otherwise totally degraded and overgrazed habitat contributes
significantly to its survival.
Taxonomy
Classification
| Class: | MAMMALIA |
| Supercohort: | LAURASIATHERIA |
| Cohort: | FERUNGULATA |
| Superorder: | CETARTIODACTYLA |
| Order: | RUMINANTIA |
| Suborder: | PECORA |
| Superfamily: | Bovoidea |
| Family: | Bovidae |
| Subfamily: | Bovinae |
| Tribe: | Neotragini (Dwarf antelope) |
| Genus: | Raphicerus |
| Species: | campestris |
The genus includes three species and two subspecies namely:
- Raphicerus campestris, the steenbok
- Raphicerus melanotis, the Cape grysbok
- Raphicerus sharpei sharpei, Sharp’s grysbok
- Raphicerus sharpei colonicus, the tropical grysbok
Former authors have
recognised 12 sub-species of steenbok these being Raphicerus campestris
bourguii, R.c. cunenensis, R.c. hoamibensis, R.c. horstockii, R.c.
kelleni, R.c. natalensis, R.c. neumanni, R.c. steinhardti, R.c.
stigmatus, R.c. tragulus, R.c. ugabensis and R.c. zukowskyi. Only six
of these occur in the southern African sub-region. Taxonomic revision
of the different sub-species is long overdue as there are only minute
distinguishable differences between them.
Image gallery
Click here to view more photographs.
Description
It is a small gracious
antelope with relatively long legs and a slender body. The skin is
smooth, pale red-brown on the back and the belly and inner thighs a
contrasting bright white. A darker brown, Y-shaped marking can
sometimes be seen on the forehead. Ewes have four teats. Adult body
size is reached within 14 to 20 months of age.
Comparison To Man

Trophy
Only rams bear fully
developed horns. These are simple, straight and spaced widely apart.
The tips are incredibly sharp and slightly bended forward. Most of the
length is smooth except for two or three rings directly above the base.
Ewes with deformed and inferior horns do occur.
Habitat requirement
Open and gently
undulating plains with scattered trees and shrubs and a short to medium
height (2-12 cm) grass layer of sweet, palatable, highly digestible
grass species are favoured. Sourveld is completely avoided. The most
suitable veld types occur within semi-arid savannahs, succulent Karoo
and grasslands. The steenbok usually avoids rocky slopes, steep
mountains, rugged country, sand-deserts, thickets and forests. Climatic
conditions can vary from semi-arid (less than 200 mm annual rainfall)
in the Kalahari, to sub-tropical (500-1 200 mm rainfall), with extreme
temperatures of -10 degrees C to 50 degrees C. Patches of tall grass
and scattered shrubs should be available as cover for protection.
Disturbed areas such as roadsides, firebreaks, landing strips, the
edges of cultivated lands and overgrazed patches are mostly favoured
due to the abundance of pioneer, broadleaf forbs. Steenbok are totally
independent of drinking water.
Optimum vegetation structure is the following canopy foliage cover:
- 40-80% cover between a height of 0 and15 cm
- 20-30% cover between 15 and 30 cm
- 15% cover between 30 and 45 cm
- 5% cover between 45 and 75 cm
- <5% cover above 75 cm.
Distribution

Feeding & Nutrition
The steenbok is a
ruminant and a mixed feeder of both broadleaf forbs and short sweet
palatable grasses of less than 12 cm that are low in fibre content. In
the Kruger National Park its diet consists of 155 dicot forb species,
42 tree and shrub species, 15 creeper species and 20 grass species. It
is a highly selective concentrate feeder with a definite preference for
specific, most delicate, parts of plants. The feeding behaviour is
dictated to by a high metabolic rate which requires a high protein and
soluble carbohydrate intake.
In general the diet consists 67% of dicot forb browse, 19% tree and
shrub browse and 6% creeper browse and only 8% grass. However, studies
on captive animals indicate that they prefer to browse exclusively,
although in some habitats grass can form up to 50% of the diet.
Steenbok are also known to dig for underground bulbs and succulent
roots, especially in semi-arid environments. They drop down on the
knees to feed at ground level and are strongly attracted to new growth
on recently burnt veld.
The mean feeding height for the steenbok is 21 cm, ranging from 0-53
cm. They are diurnal and remain active until 2-3 hours after sunset
when they lie down under thicket, in tall grass patches or in old
aardvark burrows. On moonlight nights feeding may extend later into the
dark. Growing human disturbance has caused them to become increasingly
nocturnal.
Social structure
Steenbok are mostly
solitary or occur in male/female or mother/lamb pairs. They are
strictly territorial, with the adult ram and ewe each having a
separate, fixed territory. Frequent sightings of the same individuals
in pairs in the same vicinity are a result of temporary meetings when
individuals pass through the overlaps of adjacent home ranges. Such
incidents can last up to three days and are frequently repeated. Pairs
frequently form between the same individuals but only for courtship and
mating purposes or as a mother with her lamb The common rapid
development of horns in male lambs also contributes to the
misconception of pair-bonding, as the ewe is seen in close association
with her son for an extended period giving a false impression of adult
pair-bonding.
Information Table
Steenbok information table
|
Characteristic
|
Ram
|
Ewe
|
Adult body weight
|
kg
|
9-13
|
10-13.5
|
Adult shoulder height
|
cm
|
43-50
|
45-54
|
Expected longevity
|
years
|
9
|
6-8
|
Age of sexual maturity
|
months
|
9
|
6-7
|
Age of social adulthood (1st mating)
|
months
|
13-18
|
7-9
|
Gestation
|
days
|
|
165-175
|
1st lamb born at
|
years
|
|
13-15
|
Lambing interval
|
months
|
|
8-10
|
Post maturity age (last mating)
|
years
|
-
|
-
|
Rutting season
|
Year round (peak
Feb-May)
|
| Lambing season:
|
|
Year round
(peak Sep-Dec)
|
| Weaning age |
months
|
3-4
|
Gender ratio: natural (all ages)
|
2
|
1
|
Gender ratio: production (all ages)
|
1
|
1
|
Mating ratio: natural (adults)
|
1
|
1
|
Mating ratio: production (adults)
|
1
|
1.5
|
Re-establishment: absolute minimum number needed
|
2
|
2
|
Re-establishment: smallest viable population size
|
3
|
3
|
Spatial behaviour: home range
|
ha
|
12-30
|
12-30
|
Spatial Behaviour: territory range
|
ha
|
3-15
|
5-15
|
Large stock grazing unit (adult):
Dietary ratio (grass): |
LSU
|
0.06 per animal
(34% Of diet)
|
0.06 per animal
(34% of diet)
|
Browsing unit (adult):
Dietary ratio: (browse):
|
BU
|
1.14 per animal
(66% of diet)
|
1.14 per animal
(66% of diet)
|
Maximum stocking load
|
80 animals per 1000
ha (At 400-800 mm annual rainfall)
|
Minimum habitat size required
|
ha
|
10
|
| Annual population growth |
21-32% (mean 27%) |
Optimal annual rainfall
|
300-600 mm
|
Optimal vegetation structure:
Grass height:
Woody canopy cover:
|
0-12 cm
0-40%
|
Bibliography
- Cloete, G, 1983. Etho-ecological aspects of the steenbok in the
Namib desert, South West Africa. M.SC. thesis, University of the Orange
Free State.
- Cloete, G & Kok, OB, 1986. Aspects of the water economy of steenbok in the Namib desert. Madoqua 14:375-387.
- Cloete, G & Kok, OB, 1990. Aspects of the behaviour of steenbok
in the Kuiseb River Canyon, S.W.A./Namibia. J. Nam. Sci. Soc. 42:25-46.
- Cohen, M, 1987. Aspects of the biology and behaviour of the
steenbok in the Kruger National Park. D.Sc. thesis, University of
Pretoria.
- Du Plessis, SF, 1969. The past and present geographical
distribution of the Perrisodactyla and Artiodactyla in Southern Africa.
M.Sc. Thesis, University of Pretoria.
- Du Toit, JT, 1993. The feeding ecology of a very small ruminant, the steenbok. Afr. J. Ecol. 31:35-48.
- Du Toit, JT, 1990. Home range-body mass relations: a field study on African browsing ruminants. Oecologia 85:301-303.
- Furstenburg, D 2003. Steenbok Rhapicerus campestris. Game & Hunt 9(3):6-11.
- Furstenburg, D, 2005. The Steenbok. In: Intensive Wildlife
Production in Southern Africa, Eds. Bothma, J Du P & N Van Rooyen.
Van Shaik Publishers, Pretoria, pp. 248-256.
- Haim, A & Skinner, JD, 1991. A comparative study of metabolic
rates and thermoregulation of two African antelopes, the steenbok and
the blue duiker. J. Therm. Biol. 16:145-148.
- Huntley, BJ, 1972. A note on food preferences of a steenbok. J. Sth Afr. Wildl. Mgmt Ass. 2:24-26.
- IEA (Institute of Applied Ecology), 1998. Raphiceros. In African
Mammals Databank - A Databank for the Conservation and Management of
the African Mammals Vol 1 & 2. European Commission Directorate,
Bruxelles.
- IUCN, 2006. IUCN Red list of Threatened Species, Gland, Switzerland.
- Kingdon, J, 1997. The Kingdon Field Guide to African Mammals. Princeton University Press, Princeton.
- Nowak, RM, 1991. Walker's Mammals of the World 5th edn. The Johns Hopkins University Press, Baltimore.
- Robinson, MD, 1977. An observation on parental care of young in the steenbok in South West Africa. Madoqua 10:215-216.
- Skead, CJ, 1987. Historical Mammal incidence in the Cape, Vol 1 & 2, Government Printer, Cape Town.
- Skinner, JD & Chimba, CT, 2005. The Mammals of the Southern African Subregion, 3rd edn. Cambridge University Press.
- Ward, R, 2006. Rowland Ward’s Records of Big Game, 27tth edn. Rowland Ward Publications.
- Wikipedia Encyclopedia, 2008. Connochaetus.
- Wilson, VJ & Kerr, MA, 1969. Brief notes on reproduction in steenbok. Arnoldia Rhod. 4(23):1-5.
- Wilson, DE & Reeder, DM, 1993. Mammal Species of the World: A
Taxonomic and Geographic Reference. 2nd edn., Smithsonian Institution
Press, Washington.