Kudu “Greater Kudu"
Tragelaphus strepsiceros (Pallas, 1766)

Photo: Doug Lee
| Arikaans: | Koedoe |
| German: | Kudu |
| French: | Koudou |
| Swahili: | Tandala |
| isiNdebele: | Ibhalabhala |
| isiZulu: | Umgankla/Igogo |
| isiXhosa: | Igudi |
| seSotho: | Tholo |
| seTswana: | Tholo |
| Shona: | Nhoro |
| Shangaan: | Hlongo |
| Nama: | Xaib |
| Khoi-khoi: | Ku::du |
IUCN Conservation Status:
Lower Risk, conservation dependent (LR/cd)
Kudu first became known through Kolbe’s book on “De Kaap de Goede Hoop”
in 1727. Evidence exists for the occurrence of “koo-doos” in the city
of Cape Town at the time of European colonization. The name kudu
originate from the Hottentot or Khoi-khoi word “ku::du”.
Taxonomy
Classification
| Supercohort: | LAURASIATHERIA |
| Cohort: | FERUNGULATA |
| Order: | RUMINANTIA |
| Superfamily: | BOVOIDEA |
| Family: | BOVIDAE |
| Sub-family: | Bovinae |
| Tribe: | Tragelaphini |
| Genus: | Tragelaphus |
| Species: | melampus |
There are two species:
- the greater kudu Tragelaphus strepsiceros
- the lesser kudu Tragelaphus imberbis,
and three sub-species:
- the southern greater kudu Tragelaphus strepsiceros strepsiceros
- the East African greater kudu Tragelaphus strepsiceros bea
- the northern greater kudu Tragelaphus strepsiceros cottoni
During the 1700’s and
1800’s the Eastern Cape kudu became isolated from the rest of South
Africa’s populations as a result of human settlement. At present this
population is managed as a sub-population that differs in size and
trophy quality. This is a mistake, as a genetically new sub-species is
being created artificially.
Image gallery
Click here to view more photographs.
Description
A large, slender, elegant
antelope. Adult bulls of the greater kudu are generally 35% taller than
the lesser kudu. Both sexes of the greater kudu have a mane that
continues as a whitish dorsal crest. The lesser kudu does not have a
mane. The greater kudu has 9-10 vertical white stripes, the eastern
African greater kudu 6-8 and the northern greater kudu, 4-7. The colour
of the coat differs, being a pale greyish in the southern greater kudu,
a comparatively richer colour in the eastern African greater kudu and
much paler in the northern greater kudu. The underside of the short,
furry, bushy tail flashes white when the animal is in flight. Ears are
large, round and with a white fringe. The maximum mass for cows is
reached at 4-5 years and then decreases slightly with age. Bulls do not
reach their maximum body size before the age of 12 years.
Comparison To Man

Trophy
The magnificent horns are
spread in beautiful open spirals, and smooth without grooves. The
number of turns of the spirals is related to age. There is no
scientific proof for claims that narrow horns relate to bush dwelling
or montane kudu and wide horns to plains kudu, as the two forms are
found in both habitats.
Well developed horns in bulls but cows with inferior, rudimentary,
deformed horns occasionally occur. Rowland Ward trophy status is
reached after seven years.
Habitat requirement
Broken bushveld, savannah and open woodland of deciduous plants with
scattered thicket bush clumps for refuge, both on plains and mountain
slopes. Kudu are widespread in the savannahs of eastern and southern
Africa, ranging from Sudan and Ethiopia in the north, to the Western
and Eastern Cape in the south. Their use of a specific habitat is
reliant on the density of woody plants. Kudu are seldom found in
completely open country although they may be temporarily attracted to
it by forage such as broadleaf forbs and dwarf succulents. It is
essential that the habitat contains a high diversity of fodder plants,
especially trees and shrubs, as they do not thrive on homogenous
vegetation of low diversity. Highly dense coastal dune thickets and
evergreen forests are totally avoided. Optimal annual rainfall is
300-500 mm. High mortalities are common when sudden wet, cold spells
occur, especially during periods of drought. Such mortalities were
widespread in the Karoo and Eastern Cape in 1979, 1983, 1991-’92, 1996
and 2002, most deaths being adult cows aged over six years. Kudu are
naturally diurnal but human disturbance has forced them to become
predominantly nocturnal.
Distribution

Feeding & Nutrition
Kudu are non-selective,
bulk browsers, feeding on leaves, shoots, pods or fruit of a wide range
of shrubs, trees, dicot forbs and succulents. They consume large
quantities of roughage material. In the Eastern Cape valley bushveld
the diet consists of 5-12% grass, 15-18% dicot broad leaf forbs and
70-80% tree and shrub browse. The forb to browse ratio differs greatly
with varying rainfalls and seasons. Studies in the northern savannah
mixed bushveld indicated a diet of 18% grass, 21% forbs and 61% browse.
An adult, non-lactating kudu of 210 kg consumes 3.7 kg dried plant
material per day in a dry winter and 5 kg per day in a wet summer.
There is no particular selection of young fast-fermenting plant parts,
the mean bite size measuring 3.7-4.5 cm from both old and young twig
ends. A dietary protein intake of 9-11% and 19-23% fibre should be
maintained throughout the year. Kudu can adapt to a gentle, slow change
of climate and veld condition but are intolerant of rapid changes in
food quality. A daily water intake of 7-9 litres is required in the
warmer northern and western distribution ranges.
Social structure
Kudu are semi-gregarious
and family bonding is weak. Group structures are unstable as members
constantly drift between adjacent family groups. The mean group size is
4.5, with a maximum of 20-35. During droughts temporary gatherings of
up to 60 animals can be found on open “brak”-veld. The mean number of
groups overlapping and sharing the same home range area is eight and,
areas that contain a high level of mineral salts. Kudu do not migrate
and have a fixed permanent home range which enlarge to double the size
in dry winter, but shrink again at the onslaught of a wetter summer.
Some sub-adults leave their groups and may travel up to 80 km to
establish their own home ranges. Home ranges overlap by as much as 80%
and the overlap may be shared by up to eight different groups.
The social structure comprises of:
- Family breeding groups of 1-2 socially mature bulls, 2-4 adult cows and 1-3 youngsters
- Bachelor groups of 2-6 sub-adult bulls of 2-5 years
- Socially mature male groups of 2-4 bulls of 5-8 years
- Post-mature, non-breeding male groups of 2-6 bulls over eight years
Composition of a natural population is:
- 47% socially mature adult cows aged 3-9 years
- 7% second year heifers aged >2-3 years
- 7% first-year heifers aged >1-2 years
- 18% socially mature adult bulls aged 2<8 years
- 4% post-mature trophy bulls of 8 years and older
- 9% male calves aged <1 year
- 8% female calves aged <1 year
Information Table
Greater Kudu information table
|
Characteristic
|
Bull
|
Cow
|
Adult body weight
|
kg
|
174-315
(avg. 235)
|
110-210
(avg. 155)
|
Adult shoulder height
|
cm
|
128-152
|
119-141
|
Expected longevity
|
years
|
12-16
|
7-9
|
Age of sexual maturity
|
months
|
21-24
|
15-19
|
Age of social adulthood (1st mating)
|
years
|
5
|
3
|
Gestation
|
days
|
|
250-260
|
1st calf born at
|
months
|
|
3.8-4.5
|
Calving interval
|
months
|
|
10-15
|
Post maturity age (last mating)
|
years
|
9
|
9
|
Rutting season
|
Apr-Jun
|
| Calving season:
|
|
Dec-May
|
| Weaning age |
days
|
135-165
|
Gender ratio: natural (all ages)
|
1
|
1.1
|
Gender ratio: production (all ages)
|
1
|
1.6
|
Mating
Ratio: natural (adults)
|
1
|
1.4-1.8
|
Mating
Ratio: production (adults)
|
1
|
2.5-4.2
|
Re-establishment: absolute minimum number needed
|
1
|
2
|
Re-establishment: smallest viable population size
|
3
|
9
|
Spatial behaviour: home range
|
ha
|
90-600
|
90-600
|
Spatial Behaviour: Territory
Range
|
ha
|
None
|
None
|
Large stock grazing unit (adult):
Dietary ratio (grass):
|
LSU
|
0.45 per animal
(12% of diet)
|
0.42 per animal
(12% of diet)
|
Browsing unit (adult):
Dietary ratio: (browse):
|
BU
|
1.1 per animal
(88% of diet)
|
1.1 per animal
(88% of diet)
|
Maximum stocking load
|
80 animals per 1000 ha
(at 350-450 mm annual rainfall)
|
Minimum habitat size required
|
ha
|
300
|
| Annual population growth |
13-28%
(mean 19%) |
Optimal annual rainfall
|
300-500 mm
|
Optimal vegetation structure:
Grass height:
Woody canopy cover:
|
6-6.5 cm
15-85%
|
Bibliography
- Allen-Rowlandson, TS, 1980. The social and spatial organization of
the greater kudu in the Andries Vosloo Kudu Reserve. M.Sc. Thesis,
Rhodes Univ.
- Boomker, EA, 1987. Fermentation and digestion in the kudu. D.Sc. Thesis, Univ. of Pretoria.
- Du Plessis, SF, 1969. The past and present geographical
distribution of the Perrisodactyla and Artiodactyla in Southern Africa.
M.Sc. Thesis, Univ. of Pretoria.
- Furstenburg, D, 2000. Integrated kudu, duiker, bushbuck and boer
goat production systems in Valley Bushveld: ecological interactions,
processes & constraints. Pelea 19:134-141.
- Furstenburg, D, 2002. Kudu. Game& Hunt 8(3).
- Furstenburg, D, 2005. The Kudu. In: Intensive Wildlife Production
in Southern Africa, Eds. Bothma, J Du P & N Van Rooyen. Van Shaik
Publishers, Pretoria.
- IEA (Institute of Applied Ecology), 1998. Tragelaphus. In: African
Mammals Databank - A Databank for the Conservation and Management of
the African Mammals, Vol 1 & 2. European Commission Directorate,
Bruxelles.
- IUCN, 2006. IUCN Red List of Threatened Species. Gland, Switzerland.
- Kingdon, J, 1989. East African Mammals; An atlas of evolution in Africa – Bovids, Vol 111D, Univ. of Chicago Press, Chicago.
- Novellie, PA, 1983. Feeding ecology of the kudu in the Kruger Nationasl Park. D.Sc. Thesis, Univ. of Pretoria.
- Nowak, RM, 1999. Walker's Mammals of the World 6th edn. Johns Hopkins University Press, Baltimore.
- Owen-Smith, RN, 1990. Demography of a large herbivore, the greater kudu in relation to rainfall. J. Anim. Ecol. 59:893-913.
- Owen-Smith, RN, 1994. Foraging responses of kudu to seasonal
changes in food resources: elasticity in constraints. Ecology
75:1050-1062.
- Simpson, CD, 1966. Tooth eruption, growth and ageing criteria in greater kudu. Anoldia 2:1-12.
- Simpson, CD, 1968. Reproduction and population structure in greater kudu in Rhodesia. J. Wildl. Mgmt. 32:149-162.
- Simpson, CD, 1972. Some characteristics of tragelaphine horn growth
and their relationship to age in grearter kudu and bushbuck. J. Sth.
Afr. Wildl. Mgmt. Ass. 2:1-8.
- Simpson, CD, 1972. An evaluation of seasonal movement in greater
kudu populations in three localities in South Africa. Zool. Afr.
7:197-205.
- Skead, CJ, 1987. Historical Mammal Incidence in the Cape. Vol 1 & 2, Government Printer, Cape Town.
- Skinner, JD, & Chimba CT, 2005. The Mammals of the Southern
African Subregion, 3rd edn. Cambridge University Press, Cambridge.
- Smithers, RHN, 1983. The Mammals of the Southern African Subregion,
1st edn. University of Pretoria, CTP Book Printers, Cape Town.
- Van Hoven, W, 1991. Mortalities in kudu populations related to chemical defence in trees. J. Afr. Zool. 105:141-145.
- Ward, R, 2006. Rowland Ward’s Records of Big Game, 27th edn. Rowland Ward Publications, Johannesburg.
- Wikipedia Encyclopedia, 2008. Aepyceros melampus. http://en.wikipedia.org.
- Wilson, DE & Reeder, DM, 1993. Mammal Species of the World, 2nd edn. Smithsonian Institution Press, Washington.