Impala
Aepyceros melampus (Lichtenstein, 1812)

Photo: Doug Lee
| Afrikaans: | Rooibok |
| German: | Impala |
| French: | Impala |
| isiNdebele: | Ipala / Impala |
| isiZulu: | iMpala |
| isiXhosa: | Impala |
| seSotho: | Phala |
| seTswana: | Phala |
| Shona: | Mhara |
| Shangaan: | Mbala |
| Nama: | Arab |
The name impala comes from the Zulu name iMpala while the Afrikaans name “rooibok” refers to its brick-red skin colour.
IUCN Conservation Status:
Southern & East African impala = Low Risk, conservation dependent (LR/cd)
Black-faced impala = Vulnerable (VU)
Taxonomy
Classification
| Supercohort: | LAURASIATHERIA |
| Cohort: | FERUNGULATA |
| Order: | RUMINANTIA |
| Superfamily: | BOVOIDEA |
| Family: | BOVIDAE |
| Sub-family: | Antilopinae |
| Tribe: | Aepycerotini |
| Genus: | Aepyceros |
| Species: | melampus |
Seven recognised sub-species, only three of which are accepted by the Rowland Ward trophy register:
- Southern impala Aepyceros melampus melampus including the former sub-species A.m. johnstoni, A.m. katangae and A.m. holubi
- East African impala Aepyceros.melampus suara from central, eastern Africa including the former sub-species A.m. rendilis
- Black-faced impala Aepyceros melampus petersi from northern Namibia, southern Angola and north-western Botswana
The basic body form remained almost unchanged since the Miocene (6.5
million years BP) and there has never been more than one species at any
given time. The body form bears a resemblance to many other bovids and
taxonomists consider it to be the archetypical antelope.
Image gallery
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Description
A medium-sized, lightly built antelope. The coat is shiny, uniform
chestnut brown on the back and around the neck, passing through to a
light fawn on the flanks and outer leg surfaces. The belly and the
inside of the legs are white. There is a prominent vertical black
stripe down the centre of each buttock and on the dorsal surface of the
tail. Black-faced impala are darker coloured, being a dull purple brown
with a distinctive purplish-black blaze stretching down the middle of
the face. East African impala are a brighter tawny red and are more
sharply edged along the flanks.
Comparison To Man

Trophy
Lyrate-shaped, over 50 cm
long and coarsely grooved for 75% of the length. The number of grooves
relates to the animal’s age. Well developed horns in rams but ewes with
inferior, rudimentary, deformed horns occasionally occur. Rowland Ward
trophy status is reached after five years. Black-faced impala have
smaller horns whereas those of the East African impala are longer with
a distinctive greater tip-to-tip spread.
Habitat requirement
Bushveld, savannah, open
woodland, mainly on alluvial and volcanic clay soils with an annual
mean rainfall of 400 to 700 mm. Habitats with a diverse tree and shrub
composition are favoured. Rocky outcrops, mountain slopes, open
grasslands, marshlands, arid environments, riverine thickets and
forests are avoided. Ecotones on the perimeter of riverine thickets and
closed woodland are popular. The southern impala benefits from general
increases in woodyfication due to global warming, and from overgrazing
that increases the number of annual forbs in most woodland vegetation
types. Black-faced impala prefer a denser habitat such as the riverine
thickets bordering woodland vegetation.
Distribution

Feeding & Nutrition
Mixed, concentrate feeder
and a ruminant, of both browse and short (less than 8 cm), sweet grass.
Highly selective in choice of both plant species and plant parts. A low
crude-fibre diet of less than 40% is required, together with high
protein content of 8% in winter and 16% in summer. During moist
conditions grass forms 79-92% of the diet, together with herbs, dicot
forbes and little woody browse. In dry winter the lignin content of
grass increases and the diet changes to 32-67% woody browse. Fallen
pods are an important source of stored protein during the dry winter
months.

Photo: Doug Lee
Social structure
Impala are social,
gregarious animals that stay in groups of 6-30 in a moist summer but
congregate in larger herds of 400 or more in the dry winter months
following the mating season.
- Family/breeding herds with several ewes of all ages, lambs, sub-adult rams and a few adult non-territorial rams
- Bachelor herds that form during the rut between January and April and consist of sexually mature, but socially immature rams
- Temporary nursing groups of youngsters that form on the outskirts of the family herds and are accompanied by 1-2 adult ewes
- Herds of post-mature rams of 2-4 years that are mostly of trophy quality
Information Table
Impala information table
|
Characteristic
|
Ram
|
Ewe
|
Adult body weight
|
kg
|
57 (48-65)
|
41 (38-52)
|
Adult shoulder height
|
cm
|
85-95
|
80-88
|
Expected longevity
|
years
|
10-12
|
10-12
|
Age of sexual maturity
|
months
|
16
|
13
|
Age of social adulthood (1st mating)
|
years
|
3
|
1.5
|
Gestation
|
months
|
|
6.5
|
1st lamb born at
|
months
|
|
23
|
Lambing interval
|
months
|
|
12-16
|
Post maturity age (last mating)
|
years
|
-
|
-
|
Rutting season
|
Apr-Jun
|
Lambing season:
South Africa:
Zimbabwe
East Central Africa:
Black Faced Impala:
|
|
Oct-Dec
Nov-Jan
Entire year
Dec-Jan
|
| Weaning age |
months
|
3
|
Gender ratio: natural (all ages)
|
1
|
1.1
|
Gender ratio: production (all ages)
|
1
|
1.6
|
Mating ratio: natural (adults)
|
1
|
3
|
Mating ratio: production (adults)
|
1
|
5
|
Re-establishment: absolute minimum number needed
|
2
|
3
|
Re-establishment: smallest viable population size
|
5
|
12
|
Spatial behaviour: home Range
|
ha
|
200-400
|
250-700
|
Spatial Behaviour: territory range
|
ha
|
4-10
|
None
|
Large stock grazing unit (adult):
Dietary ratio (grass):
|
LSU
|
0.17 per animal
(45% of diet)
|
0.17 per animal
(45% of diet)
|
Browsing unit (adult):
Dietary ratio: (browse):
|
BU
|
0.4 per animal
(55% of diet)
|
0.4 per animal
(55% of diet)
|
Maximum stocking load
|
300 animals per 1000 ha
(at 350-450 mm annual rainfall)
|
Minimum habitat size required
|
ha
|
150
|
Annual population growth
|
23-48%
(mean 35%)
|
Bibliography
- Anderson, JL, 1971. Seasonal changes in the social organization and
distribution of the impala in Hluhluwe game reserve, Zululand. J. sth
Afr. Wildl. Mgmt. Ass. 2:16-20.
- Brooks, PM, 1978. Relationship between body condition and age,
growth, reproduction and social status in impala, and its application
for management. Sth. Afr. J. Wildl. Res. 8(4):151-157.
- Child, G, 1964. Growth and ageing criteria of impala Aepyceros melampus. Arnoldia 27:128-135.
- Du Plessis, SF, 1969. The past and present geographical
distribution of the Perrisodactyla and Artiodactyla in Southern Africa.
M.Sc. Thesis, University of Pretoria.
- Estes, RD, 1991. The behaviour guide to African mammals including
hoofed mammals, carnivores, primates. University of California Press,
California.
- Fairall, N & Braack, HH, 1976. Growth and development of the impala Aypyceros melampus. Koedoe 19:83-88.
- Furstenburg, D, 1970-2008. Personal field notes (unpublished).
- Furstenburg, D, 1997. Rooibok. S.A. Wild & Jag 3(4).
- Furstenburg, D, 2006. Rooibok Aepyceros melampus. Wild & Jag 12(7).
- Furstenburg, D, 1984. Die voedingsekologie van die rooibok
Aepyceros melampus as deel van die Savanna Ekosisteem Projek, CSIR,
Nyl;svley Natuurreservaat. Unpublished report, Ugené Marais Chair in
Wildlife Management, University of Pretoria.
- Furstenburg, D, 2005. The Impala. In: Intensive Wildlife Production
in Southern Africa, Eds. Bothma, J Du P & N Van Rooyen. Van Shaik
Publishers, Pretoria.
- Green, WC & Rothstein, A, 1998. Translocation, hybridization
and the endangered black-faced impala. Conserv. Biol. 12:475-480.
- Hill, RH, 1982. Effect of dietary extreme of impala rumen epimural flora. Appl. Enviro. Microbiol. 44:198-202.
- Howewls, WW & Hanks, J, 1975. Body growth of the impala
Aepyceros melampus in Wankie National Park, Rhodesia. J. sth Afr.
Wildl. Mgmt. Ass. 5(2):95-98.
- IEA (Institute of Applied Ecology), 1998. Aepyceros melampus. In:
African Mammals Databank - A Databank for the Conservation and
Management of the African Mammals, Vol 1 & 2. European Commission
Directorate, Bruxelles.
- IUCN, 2006. IUCN Red List of Threatened Species. Gland, Switzerland.
- Jarman, MV & Jarman, PJ, 1973. Daily activity of impala. East Afr. Wildl. J. 11:75-92.
- Joubert, E, 1971. Observations on the habitat preferences and
populatiuon dynamics of the black-faced impala Aepyceros petersi in
S.W. Africa. Madoqua 1:55-65.
- Kingdon, J, 1997. The Kingdon Field Guide to African Mammals. Princeton University Press, Princeton
- Kingdon, J, 1989. East African Mammals; An atlas of evolution in
Africa – Bovids, Vol 111D, University of Chicago Press, Chicago.
- Meissner, HH, Pieterse, E & Potgieter, JHJ, 1996. Seasonal food
selection and intake by male impala in two habitats. Sth. Afr. J.
Wildl. Res. 26:56-63.
- Monro, RH, 1979. A study on the growth, feeding and body condition of impala. M.Sc. dissertation, Pretoria University, Pretoria.
- Murray, MG, 1980. Social structure of an impala population. D.Phil. thesis, University of Rhodesia.
- Murray, MG, 1982. Home range, dispersal and the clan system of impala. Afr. J. Ecol. 20:253-369.
- Nowak, RM, 1999. Walker's Mammals of the World 6th edn. Johns Hopkins University Press, Baltimore.
- Opperman, RB, 1983. Feeding ecology of the impala in Northern Transvaal. M.Sc. dissertation, Pretoria University, Pretoria.
- Skead, CJ, 1987. Historical Mammal Incidence in the Cape. Vol 1 & 2, Government Printer, Cape Town.
- Skinner, JD, & Chimba CT, 2005. The Mammals of the Southern
African Subregion, 3rd edn. Cambridge University Press, Cambridge.
- Smithers, RHN, 1983. The Mammals of the Southern African Subregion,
1st edn. University of Pretoria, CTP Book Printers, Cape Town.
- Van Rooyen, AF, 1996. Simulated protein requirements and seasonal
breeding in impala Aepyceros melampus. Sth. Afr,. J. Wildl. Res.
26:77-80.
- Vincent, J, 1972. Reproduction in the impala with particular
reference to the population in the Mkuze Game reserve, Zululand. M.Sc.
thesis, Pretoria University, Pretoria.
- Vincent, J, 1979. The population dynamics of impala in the Mkuze Game reserve, Zululand. Ph.D. thesis, University of Natal.
- Ward, R, 2006. Rowland Ward’s Records of Big Game, 27th edn. Rowland Ward Publications, Johannesburg.
- Wikipedia Encyclopedia, 2008. Aepyceros melampus. http://en.wikipedia.org.
- Wilson, DE & Reeder, DM, 1993. Mammal Species of the World, 2nd edn. Smithsonian Institution Press, Washington.
- Young, E, 1972. Observations on the movement patterns and daily
range size of impala in the Kruger National Park. Zool. Africana
7:187-195.