Black Wildebeest / White-tailed gnu
Connochaetes gnou (Zimmermann, 1780)

Photo: Sam Bash
| Afrikaans: | Swartwildebees |
| German: | Weisschwanz gnu |
| French: | Gnou à queue blanche |
| isiZulu: | Inkonkoni |
| isiXhosa: | Inqu |
| Sesotho: | Pudumo |
| Tswana: | Pudomô |
| Shona: | Mbutuma |
| Khoi-khoi: | Gnu |
IUCN Conservation Status:
Lower Risk, least concern (LR/lc)
This odd animal is certainly the most bizarre living antelope with the
mane and tail of a horse, the face of a steer and the delicate legs of
a buck. Its appearance is matched by its freaky behaviour. A phrase by
W Cornwallis Harris (1840), read as follows, “A more whimsical compound
than the Gnoo could scarcely have been thrown together, or a monster
imagined of more fantastical and anomalous exterior”. The name ‘black”
is misleading as the animal is a dark brown, while the name
“white-tailed gnou”, which refers to the cream-white tail, is more
relevant. “Gnu” is a Khoi-khoi word named after the animals’ alarm
snort, a “ger-nu”.
Taxonomy
Classification
| Class: | MAMMALIA |
| Supercohort: | LAURASIATHERIA |
| Cohort: | FERUNGULATA |
| Superorder: | CETARTIODACTYLA |
| Order: | RUMINANTIA |
| Suborder: | PECORA |
| Superfamily: | Bovoidea |
| Family: | Bovidae |
| Subfamily: | Bovinae |
| Tribe: | Alcelaphini |
| Genus: | Connochaetes |
| Species: | gnou |
The tribe Alcelaphini
evolved in Africa about 6 million years BP and includes the three
genera Connochaetus or wildebeest, Damaliscus the blesbok and tsessebe
and Alcelaphus the hartebeest. There are two species and four
subspecies of wildebeest
- Connochaetes gnou the black wildebeest
- Connochaetes taurinus the blue wildebeest; subspecies
- C.t. taurinus the southern brindled gnu (blue wildebeest)
- C.t. cooksoni the Nyassa or Luangwa valley brindled gnu
- C.t. johnstoni the Mozambique brindled gnu
- C.t. albojubatus the East African white-bearded gnu.
The
black wildebeest is endemic to South Africa and never existed
elsewhere. Hundreds of thousands once roamed the open grasslands of the
Free State and the Northern Cape Provinces, and lesser numbers in the
central grasslands of most other provinces.
Description
A large antelope with a
stiff, straight mane of long, pale-cream hair tipped with brown; a long
white, horse-like tail; and the brush of brown, curly hair on the
muzzle. The body is a dark yellow-brown that appears black from a
distance. The face is darker in colour than the body. The back is
hollow with both the shoulders and the buttocks at a higher level than
the rump. A long beard stretches from the throat across the centre of
the breast to the navel. Calves are a cream to light-brown at birth
that continues to darken until an age of 18 months.
Comparison to Man

Trophy
Both genders have
well-developed horns that have their origins in a thickened boss on the
head and then extend forwards in a deep, downwards curve. The anterior
halves of the horns are straight and then project vertically upwards
with the tips bent slightly backwards. The horns of cows are thinner
than those of the bulls and are thus lighter. Horn development is
relatively rapid and trophy status may be reached at 2.5 years of age.
Habitat requirement
Open grassy plains, with
short grasses and without trees and shrubs are preferred. The grass
composition may be either a sweet or a mixed veld type but preferably
not exclusively sour. Highveld areas and mountain plateaux’s are
favoured, especially at an annual rainfall of 400-700 mm. Semi-arid and
desert conditions and sub-tropical or tropical habitats are unsuitable.
Forests, thickets, closed woodland, bushveld and wooded drainage lines
are also generally avoided. If located in a habitat that is not
optimal, black wildebeest will transform the structure of the
herbaceous vegetation destructively by ploughing the soil with the
horns and front hooves. Tall grass is not tolerated but new growth on
recently burnt veld is a major attraction. Surface water for daily
drinking is essential.
Distribution

Feeding & Nutrition
Most feeding takes place during the cooler daylight hours of the early
morning and late afternoon. Hot midday hours are spend lying in open
sunlight ruminating. Black wildebeest are exclusively grazers and
highly selective of both the species of grass and its vegetative part.
Artificial licks and supplementary, concentrated pellets are not easily
accepted. The natural diet consists of between 63-93% short grasses
below 6 cm of both sweet and sour species, 3-34% karroid dwarf shrubs
and brush and 3% dicot forbs. Grasses above 12 cm are marginal to
unsuitable.
Social structure
Black wildebeest are social gregarious animals. The structure of a population consists of:
• family groups of 10-60 individuals consisting of adult cows, heifers, post-mature cows and calves of both sexes
• bachelor groups of 10-30 individuals including sub-adult bulls of 1-3 years and non-dominant adult bulls
• territorial breeding bulls which are dominant and mostly solitary.
Adult cows maintain a hierarchy of female dominance that is mostly
age related. The highest ranked female acts as group leader. Family
bonds between cows and their female offspring are very tight and can
last for life. Male calves of 12-14 months are chased from the herd by
the dominant bulls as the herd moves through their territories.
Post-mature bulls cast out from the herd usually reunite with bachelor
groups. In absence of competition by developing bulls, a single
territorial bull in an area will often attack heifers and calves
aggressively out of frustration. They are area bound and will remain in
the same home range as long as the veld condition remains suitable and
sufficient, otherwise they will migrate over long distances. The adult
body size is reached at 2 years.
As black and blue
wildebeest crossbreed successfully, the two species should not be kept
together on small management units or within the same habitat. In the
wild however, the two species rarely interact due to their different
habitat preferences. A manager must be constantly aware of the
potential risk of crossbreeding. Nature Conservation authorities do not
issue permits for keeping both species on the same management unit.
Crossbreeds are known commercially as red wildebeest. Red wildebeest
hybrids must not be confused with the recent commercially bred golden
wildebeest, which is not a genetic hybrid but a colour variation of the
blue wildebeest.
Diseases
Black wildebeest are
commonly resistant to the majority of diseases. Exceptions are the
larvae of the nasal butt-flies Kirkioestrus minentus and Oestrus
variolosus, ocular-vascular myiasis caused by the larvae of the fly
Gidoelstia sp. and malignant catarrhal fever. As the latter two
diseases are highly contagious in domestic livestock, especially in
cattle, both black and blue wildebeest must be separated from them by a
corridor of at least 1 km. However malignant catarrhal fever is more
commonly found in blue wildebeest with their sub-tropical origin, than
in black wildebeest from the cooler temperate regions. Exposing black
wildebeest to either blue wildebeest or to warmer sub-tropical
conditions increases the risk of infection.
Information Table
Black Wildebeest information table
|
Characteristic
|
Bull
|
Cow
|
Adult body weight
|
kg
|
147-193
|
120-160
|
Adult shoulder height
|
cm
|
120
|
110
|
Expected longevity
|
years
|
16
|
18
|
Age of sexual maturity
|
months
|
16
|
16
|
Age of social adulthood (1st mating)
|
months
|
3
|
1.5
|
Gestation
|
days
|
|
255
|
1st calf born at
|
years
|
|
2.4
|
Furrow interval
|
months
|
|
12
|
Post maturity age (last mating)
|
years
|
-
|
-
|
Rutting season
|
Feb-Apr
|
| Calving season:
|
|
Nov-Jan
|
| Weaning age |
months
|
4-5
|
Gender ratio: natural (all ages)
|
1
|
1.8
|
Gender ratio: production (all ages)
|
1
|
4
|
Mating ratio: natural (adults)
|
1
|
3
|
Mating ratio: production (adults)
|
1
|
6-10
|
Re-establishment: absolute minimum number needed
|
2
|
3
|
Re-establishment: smallest viable population size
|
3
|
8
|
Spatial behaviour: home range
|
ha
|
400
|
200-500
|
Spatial behaviour: territory range
|
ha
|
2-6 (non static)
|
None
|
Large stock grazing unit (adult):
Dietary ratio (grass): |
LSU
|
0.35 per animal
(93% of diet)
|
0.35 per animal
(93% of diet)
|
Browsing unit (adult):
Dietary ratio: (browse):
|
BU
|
1.11 per animal
(7% of Diet)
|
1.11 per animal
(7% of Diet)
|
Maximum stocking load
|
100 animals per 1000
ha (At 450-550 mm annual Rainfall)
|
Minimum habitat size required
|
ha
|
400
|
| Annual population growth |
28-37% (mean 32%) |
Optimal annual rainfall
|
450-700 mm
|
Optimal vegetation structure:
Grass height:
Woody canopy cover:
|
2-12 cm
0-15%
|
Bibliography
- Du Plessis, SF, 1969. The past and present geographical
distribution of the Perrisodactyla and Artiodactyla in Southern Africa.
M.Sc. Thesis, University of Pretoria.
- Furstenburg, D 2007. Swartwildebees Connochaetus gnou. Wild & Jag 13(2).
- Furstenburg, D, 1970-2008. Personal field notes (unpublished).
- IEA (Institute of Applied Ecology), 1998. Connochaetus gnou. In
African Mammals Databank - A Databank for the Conservation and
Management of the African Mammals Vol 1 & 2. European Commission
Directorate, Bruxelles.
- IUCN, 2006. IUCN Red list of Threatened Species, Gland, Switzerland.
- Kingdon, J, 1997. The Kingdon Field Guide to African Mammals. Princeton University Press, Princeton.
- Kok, OB & Vrahimis, S, 1995. Black wildebeest territorial clearings. J. Afr. Zool. 109:231-237.
- Nowak, RM, 1991. Walker's Mammals of the World 5th edn. The Johns Hopkins University Press, Baltimore.
- Skead, CJ, 1987. Historical Mammal incidence in the Cape, Vol 1 & 2, Government Printer, Cape Town.
- Skinner, JD & Chimba, CT, 2005. The Mammals of the Southern African Subregion, 3rd edn. Cambridge University Press.
- Von Richter, W, 1971. Past and present distribution of the black
wildebeest, Connochaetus gnou, with special reference to the history of
some herds in South Africa. Ann. Trans. Mus. 27:35-57.
- Von Richter, W, 1971. Observations on the biology and ecology of
the black wildebeest, Connochaetus gnou. J. Sth. Afr. Wildl. Mgmt. Ass.
1:3-16.
- Von Richter, W, 1972. Territorial behaviour of the black wildebeest, Connochaetus gnou. Zool. Afr. 7:207-231.
- Vrahimis, S, 2000. Aspects of the ecology ofof black wildebeest in a semi-arid environment. Afr. J. Ecol. 30:169-175. black wildebeest. Pelea 19:88-91.
`. Vrahimis, S & Kok OB, 1992. Body orientation
- Vrahimis, S & Kok OB, 1993. Daily activity of black wildebeest in a semi-arid environment. Afr. J. Ecol. 31:328-336.
- Ward, R, 2006. Rowland Ward’s Records of Big Game, 27tth edn. Rowland Ward Publications.
- Wikipedia Encyclopedia, 2008. Connochaetus.
- Wilson, DE & Reeder, DM, 1993. Mammal Species of the World: A
Taxonomic and Geographic Reference. 2nd edn., Smithsonian Institution
Press, Washington.